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This book is about a nine year old boy named Bruno who, because of his father's job, is forced to move from Berlin to the countryside. From his bedroom window, Bruno spots a fence behind which he sees men in 'striped pyjamas'. These are Jews, and they are in a Nazi concentration camp. One day his parents come to an agreement that both Bruno and Gretel need a tutor for their education, so they hire Herr Liszt. To Bruno, Herr Liszt is the most boring teacher one could ever have; he teaches social studies instead of reading and arts, which Bruno prefers. So, in boredom and confusion, Bruno wonders what is going on at "Out-With" (Auschwitz) and why people are always dressed in striped pyjamas there. One afternoon, he goes exploring, and meets a Jewish boy called Shmuel, a name Bruno has never before heard but that apparently is quite common in the concentration camp. Shmuel soon becomes Bruno's friend and Bruno visits every afternoon to talk. Bruno is told that the people in the striped pyjamas on the other side of the fence are Jews and that he and his family are "the opposite".

The story ends with Bruno about to return to the family's old home in Berlin. As a final adventure, he agrees to dress in a set of striped pyjamas and climb under a loose wire in the fence to help Shmuel find his father, who went missing in the camp. The boys are unable to find him, and just as it starts to rain and get dark, Bruno decides he would like to go home, yet the Nazis in the area of the camp force the boys to go on a march. Neither boy knows where this march will lead. However, they are soon crowded into a gas chamber, and the author leaves the story with Bruno pondering, yet unafraid, in the dark holding hands with Shmuel. "...Despite the chaos that followed, Bruno found that he was still holding Shmuel's hand in his own and nothing in the world would have persuaded him to let go".

In an epilogue, Bruno's family spend several months at their home trying to find Bruno,

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