Arthropods are characterized by the possession of a segmented body with appendages on each segment. They have a dorsal heart and a ventral nervous system. All arthropods are covered by a hard exoskeleton that is made out of chitin, a polysaccharide. Periodically, an arthropod sheds this covering when it moults. This covering makes arthropods less prone to dehydration.
The success of the arthropods is related to their hard exoskeleton, segmentation, and jointed appendages. The appendages are used for feeding, sensory reception, defense, and locomotion. The muscle system is more or less assisted by hydraulics originated from the blood pressure created by the heart. The hydraulic system in spiders is especially well developed.
Most arthropods breathe through a tracheal system (exceptions are arthropods like sea spiders, Pauropoda, some thysanurans and some arachnids like many mites who breathe through their body surface instead); a potential difficulty considering that the skeletal structure is external and covers nearly all of the body. Aquatic arthropods use gills to exchange gases. These gills have an extensive surface area in contact with the surrounding water. Terrestrial arthropods have internal surfaces that are specialized for gas exchange. Insects and most other
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